Saturday, 5 May 2012

List the major features of a typical animal cell and a higher plant cell, respectively.


Major Features of a Typical Animal Cell
Structure  Molecular Composition Function
Extracellular matrix The surfaces of animal cells are covered with a flexible and sticky layer of complex carbohydrates, proteins, and lipids. This complex coating is cell-specific, serves in cell–cell recognition and communication, creates cell adhesion and provides a protective outer layer
Cell membrane
(plasma membrane)
Roughly 50; 50 lipid; protein as a 5-nm-thick continuous sheet of lipid bilayer in which a variety of proteins are embedded. The plasma membrane is a selectively permeable outer boundary of the cell,  containing specific systems—pumps, channels, containing specific systems—pumps, channels, transporters—for the exchange of nutrients and other materials with the environment.  Important enzymes are also located here.
Nucleus The nucleus is separated from the cytosol by  a double membrane, the nuclear envelope. The DNA is complexed with basic proteins (histones) to form chromatin fibers, the material from which chromosomes are made. A distinct RNA-rich region, the nucleolus, is the site of ribosome assembly. The nucleus is the repository of genetic information encoded in DNA and organized into chromosomes. During mitosis, the chromosomes are replicated and transmitted o the daughter cells. The genetic information of DNA is transcribed into RNA in the nucleus and passes into the cytosol where it is translated into protein by ribosomes.
Mitochondria  Mitochondria are organelles surrounded by two membranes that differ markedly in their protein and lipid composition. The inner membrane and its interior volume, the matrix, contain many important enzymes of energy metabolism. Mitochondria are about the size of bacteria, »1mm. Cells contain hundreds of mitochondria, which collectively  occupy about one-fifth of the cell volume. Mitochondria are the power plants of eukaryotic cells where carbohydrates, Fats, and amino acids are oxidized to CO2 and H2O. The energy released is trapped as high-energy phosphate bonds in ATP.  
Golgi apparatus A system of flattened membrane-bounded vesicles often stacked into a complex. Numerous small vesicles are found peripheral to the Golgi and contain secretory material packaged by the Golgi. Involved in the packaging and processing of macromolecules for secretion and for delivery to other cellular compartments.  
Endoplasmic reticulum
 (ER) and ribosomes
Flattened sacs, tubes, and sheets of internal membrane extending throughout the cytoplasm of the cell and enclosing a large  interconnecting series of volumes called cisternae. The ER membrane is continuous with the outer membrane of the nuclear envelope. Portions of the sheetlike areas of  the ER are studded with ribosomes, giving rise to rough ER. Eukaryotic ribosomes are larger than prokaryotic ribosomes. The endoplasmic reticulum is a labyrinthine organelle where both membrane proteins and lipids are synthesized. Proteins made  by the ribosomes of the rough ER pass  through the outer ER membrane into the cisternae and can be transported via the Golgi to the periphery of the cell. Other ribosomes unassociated with the ER carry on protein synthesis in the cytosol.
Lysosomes Lysosomes are vesicles 0.2–0.5 mm in diameter, bounded by a single membrane. They contain  hydrolytic enzymes such as proteases and  nucleases which, if set free, could degrade  essential cell constituents. They are  formed by budding from the Golgi apparatus. Lysosomes function in intracellular digestion of materials entering the cell via phagocytosis or pinocytosis. They also function in the controlled degradation of cellular components.
Peroxisomes Like lysosomes, peroxisomes are 0.2–0.5 mm  single-membrane–bounded vesicles. They contain a variety of oxidative enzymes that use molecular oxygen and generate peroxides. They are formed by budding from the smooth ER.  Peroxisomes act to oxidize certain nutrients, such as amino acids. In doing so, they form potentially toxic hydrogen peroxide, H2O2, and then decompose it to H2O and O2 by way of the peroxide-cleaving enzyme catalase.
Cytoskeleton The cytoskeleton is composed of a network of protein filaments: actin filaments (or microfilaments), 7 nm in diameter;  intermediate filaments, 8–10 nm; and microtubules, 25 nm. These filaments interact in establishing the structure and functions of the cytoskeleton. This interacting network of protein filaments gives structure and  organization to the cytoplasm.     The cytoskeleton determines the shape of the cell and gives it its ability to move. It also mediates the internal movements that occur in the cytoplasm, such as the migration of organelles and mitotic movements of chromosomes. The propulsion  instruments of cells—cilia and flagella—are constructed of microtubules.
     

Table 1.8
Major Features of a Higher Plant Cell: A Photosynthetic Leaf Cell
Structure                       
Molecular Composition
Function
Cell wall  Cellulose fibers embedded in a  polysaccharide/protein matrix; it is thick (á0.1 mm), rigid, and porous to small molecules. Protection against osmotic or mechanical rupture. The walls of neighboring cells interact in cementing the cells together to form the plant. Channels for fluid circulation and for cell–cell communication pass through the walls. The structural material confers form and strength on plant tissue.
Cell membrane Plant cell membranes are similar in overall structure and organization to animal cell membranes but differ in lipid and protein composition. The plasma membrane of plant cells is selectively permeable, containing transport systems for the uptake of essential nutrients and inorganic ions. A number of important enzymes are localized here.
Nucleus The nucleus, nucleolus, and nuclear envelope of plant cells are like those of animal cells. Chromosomal organization, DNA replication, transcription, ribosome synthesis, and mitosis in plant cells are grossly similar to the analogous features in animals.
Chloroplasts Plant cells contain a unique family of organelles, the plastids, of which the chloroplast is the prominent example. Chloroplasts have a double membrane envelope, an inner volume called the stroma, and an internal membrane system rich in thylakoid membranes, which enclose a third compartment, the thylakoid lumen. Chloroplasts are significantly  larger than mitochondria. Other plastids are found in specialized structures such as fruits, flower petals, and roots and have specialized roles. Chloroplasts are the site of photosynthesis, the reactions by which light energy is converted to metabolically useful chemical energy in the form of ATP. These reactions occur on the thylakoid membranes. The formation of carbohydrate from CO2 takes place in the stroma. Oxygen is evolved during photosynthesis. Chloroplasts are  the primary source of energy in the light.
Mitochondria Plant cell mitochondria resemble the mitochondria of other eukaryotes in form and function. Plant mitochondria are the main source of energy generation in photosynthetic cells in the dark and in nonphotosynthetic cells under all conditions.
Vacuole The vacuole is usually the most obvious compartment in plant cells. It is a very large vesicle enclosed by a single membrane called the  tonoplast. Vacuoles tend to be smaller in young cells, but in mature cells, they may occupy more than 50% of the cell’s volume. Vacuoles occupy the center of the cell, with the cytoplasm being located peripherally around it. They resemble the lysosomes of animal cells. Vacuoles function in transport and storage of nutrients and cellular waste products. By accumulating water, the vacuole allows the plant cell to grow dramatically in size with no increase in cytoplasmic volume.
Golgi apparatus, endoplasmic reticulum, ribosomes, lysosomes, peroxisomes, and cytoskeleton Plant cells also contain all of these characteristic eukaryotic organelles, essentially in the form described for animal cells. These organelles serve the same purposes in plant cells that they do in animal cells.
  
            Eukaryotic cells possess a discrete, membrane-bounded nucleus, the repository of the cell’s genetic material, which is distributed among a few or many chromosomes. During cell division, equivalent copies of this genetic material must be passed to both daughter cells through duplication and orderly partitioning of the chromosomes by the process known as mitosis. Like prokaryotic cells, eukaryotic cells are surrounded by a plasma membrane. Unlike prokaryotic cells, eukaryotic cells are rich in internal membranes that are differentiated into specialized structures such as the endoplasmic reticulum (ER) and the Golgi apparatus. Membranes also surround certain organelles (mitochondria and chloroplasts, for example) and various vesicles, including vacuoles, lysosomes, and peroxisomes. The common purpose of these membranous partitionings is the creation of cellular compartments that have specific, organized metabolic functions, such as the mitochondrion’s role as the principal site of cellular energy production. Eukaryotic cells also have a cytoskeleton composed of arrays of filaments that give the cell its shape and its capacity to move. Some eukaryotic cells also have long projections on their surface—cilia or flagella—which provide propulsion.

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